Biochemistry NAD+ is reduced to NADH + H+ by dehydrogenases of all the following substrates, Except A Pyruvate B Glyceraldehyde-3-phosphate C Malate D Succinate 💡 Explanation NAD+ is reduced to NADH + H+ by dehydrogenases of all the following substrates, Except ✓ Correct Answer: D. Succinate